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Plant Science Laboratories Herbarium News No. 34
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Plant Science Laboratories
Herbarium News No. 34
ISSN 0953-0080 Tuesday 1st November 1994
It is now well past the time that an issue of Herbarium News should have been produced.
In fact, there has been so much activity and news to report that even the thought of
`putting pen to paper' became rather daunting! Herbarium News started in October 1983
(although the first issue was undated) when there was a gap in PSL's own news-sheet
(Interflora). The fact that it has gone from strength to strength is the result of the
combined efforts of a number of staff and you for helping provide the information! Please
continue to send in to me items for the next Herbarium News: Dr Stephen L. Jury,
Department of Botany, Plant Science Laboratories, The University of Reading,
Whiteknights, Reading, Berkshire RG6 2AS, UK, Telephone: +44.(0)734.318169, Fax:
It is very pleasing to report that Dr Anne Bruneau and Dr Terry Hedderson are now in
post, though sadly work permits were somewhat delayed, especially in Terry's case.
I am also pleased to say that the University has also supported this taxonomy initiative by
allowing Anne and Terry each to have a postgraduate research student through the
University Research Endowment Fund. Mr Cymon Cox of Bangor is now in post to
research the phylogenetic relationships in the Bryoideae (Musci) inferred from rDNA
sequences with Terry, and our own Bill Baker, who has just completed our MSc course
(MSc project: A revision of the Cyrtorchis arcuata (Lindl.) Schltr. complex
(Orchidaceae), will be undertaking a systematic study of the Calamoid Palms. Molecular
techniques will be used to examine generic-level relationships in Calamoideae and the

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results compared with morphological and palynological data within a cladistic
As yet we have no word on the accompanying NERC postdoctoral Research Fellowships;
applications from Reading and Birmingham are still being evaluated by NERC.
The arrival of Joe Mullins from Dublin (see elsewhere under `Botanical Diversity of the
Indo-Pacific Region') to work with Dr Alastair Culham on a new molecular project,
further strengthens our Molecular Systematics group. (Most of our systematics research
students are joining this group for a period to add a molecular component to their
We wish to congratulate Michael on being given a personal chair and becoming Professor
of Mycobiology. Nobody has used this title `Mycobiology' before, but it serves to
emphasize the importance of fungi in biology!
This honour has been given in recognition of Michael's outstanding contribution to the
taxonomy and ecology of the Straminipiles. He is the world authority on this kingdom of
major importance and the distillation of his knowledge will be presented soon in a
massive publication which will be the definitive work on the subject.
September 30th saw the retirement of Professor David Moore. A traditional tea party took
place in the PSL foyer. The numerous participants, including several ex-students, heard
Professor Michael Dick outlining some of David's incredible range of achievements. Dr
Royce Longton presented a volume of letters, tributes, messages and photographs
collected together by Mrs Ida Moore from David's friends and students, past and present
(spanning 5+ decades and 6 continents). David received a large collection of Do-It-
Yourself tools (electric circular saw, power belt sander, Tenon saw, Hand saw, Hand
drill), plus Heelas vouchers and a book token. David responded by outlining the
important r“le of happenstance in the evolution of at least one botanical career. He is at
present busy writing over 150 thank- you letters!
Besides Cymon Cox and Bill Baker mentioned elsewhere, we welcome:

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Research students
Shaida Fariza Sulaiman (ex-USM, Malaysia) has started a research project entitled
`Chemotaxonomical and pharmacological studies of the Mimoseae'. Shaida has already
converted in excess of 100 herbarium specimens into phytochemical extracts.
Jose Augusto Carvalho (University of Madeira, Funchal) will be studying the
phytogeographical relationships of selected Macaronesian taxa. We are pleased to say
that Bob Press (BM) will be actively involved in this project.
Diploma in Pure and Applied Plant and Fungal
Stephen CAFFERTY Royal Botanic Gardens, Kew
Murray DAWSON Landcare Research, New Zealand
Alan HULME Royal Botanic Gardens, Kew
David NANGOMA National Herbarium & Botanic Gardens
Malawi, Malawi
MSc in Pure and Applied Plant and Fungal Taxonomy
Ben ARNOLD University of St Andrews
Sarah COURTNEY University of Liverpool
Laura FORREST University of Edinburgh
Joelle HAUZEUR Ecole Pratique des Hautes Etudes,
Wan-Pyo HONG Chungbuk National University, Korea
Yusuf MENEMEN Kirikkale University, Turkey
Momtaz MIRZA Bangladesh National Herbarium, Dhaka,
Zuzana PRICE College of Agriculture, Plovdiv,

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After an absence of seven years, during which he worked for IUCN - The World
Conservation Union as its Chief Scientist, Plant Conservation and as Director of Botanic
Gardens Conservation International, Professor Heywood has returned to the Department
of Botany. He will now be working mainly for the United Nations Environment
Programme as Executive Editor of its Global Biodiversity Assessment (GBA). The
objective of the GBA is to provide an independent, critical, peer-reviewed, scientific
analysis of the current issues, theories and views concerning the main global aspects of
biological diversity. These aspects include the characterization, origins, dynamics,
magnitude, distribution, monitoring, and multiple values of biodiversity; biodiversity and
ecosystem function; human influences on biodiversity; conservation and sustainable use
of biodiversity; biotechnology; data and information management and communication.
The project is financed by the Global Environment Facility (GEF). The Chairman of the
GBA is Dr Robert T. Watson, Office of Science and Technology of the White House, and
Honorary Advisers are Dr Peter Raven, Dr Emile Salim, Dr M.S. Swaminathan and
Professor E.O. Wilson. As Executive Editor, Professor Heywood is co-ordinating the
working of the co-ordinators and teams of authors of each of the twelve sections of the
GBA. Several hundred scientists from over 40 countries are engaged in the preparation of
the Assessment, the publication of which is expected in late 1995.
Professor Heywood is also chairing a steering group, under the IUBS/UNESCO
DIVERSITAS programme, which is developing a new theme on the "Conservation of the
genetic variability of wild species". On a related theme he is a member of the Council of
Europe Group of Specialists on Biodiversity and Biosubsistence and has been an
organizer of three workshops held in Faro (1992), Neuchatel (1993) and Palermo (1994)
on the "Conservation of the Wild Relatives of European Crop Plants" under the auspices
of the Group. He was also recently elected the first President of the IUBS International
Council of Medicinal and Aromatic Plants.
Professor Heywood is also currently finishing the editing of the IUCN/WWF project on
"Centres of Plant Diversity", the first volume of which is due to be published in
November 1994.
There was a meeting of this European Union Network on the 5th and 6th October 1994 in
the Rijksherbarium, Leiden, attended by Drs Culham and Jury. The meeting included
talks from the postgraduates and post-doctoral fellows on their research projects. The
following eight staff have been appointed to the nine nodes in the network:
1. Dr Stefan Dressler (from Germany, now in Leiden): Euphorbiaceae - Bridelia.
2. Mr Josef Mullins (from Ireland, now in Reading): Nepenthaceae - Nepenthes.
3. Dr David Middleton (from Scotland, now in Dublin): Apocynaceae.
4. Dr Petra Hoffman (from Germany, now at Kew): Euphorbiaceae - Antidesma.
5. Dr Helen Fortune-Hopkins (from England, now in Paris): Cunoniaceae

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6. Mr Rogier de Kok (from Leiden, now at Oxford): Labiatae/Verbenaceae.
7. Ms Josefine Schneidt (from Germany, now at Aberdeen): Asclepiadaceae.
8. Ms Nicoletta Cellinese (from Italy but working at BM, [a Reading graduate],
going to Mainz): Melastomataceae - Sonerila.
At the time of writing, the postgraduate position in Aarhus is still not filled. I am
informed by Dr Marco Roos in Leiden that an EC postgraduate student is needed urgently
(certainly by 1 December 1994). Please contact him directly by fax: +31.71.273511 in
Leiden. The post is for two years. The next meeting will be held at Kew in conjunction
with the Flora Malesiana symposium next July.
At Reading, Joe's study will involve production of a sectional- level classification within
a phylogenetic framework for the genus Nepenthes. The work will employ morphological
and molecular data in a cladistic analysis.
Collecting for our collaborative project `Floristic Biodiversity of Northern Morocco'
commenced again in February with Dr Stephen Jury, Mr Tim Upson (of Reading) and Mr
Graham Walters (of the Alpine and Herbaceous Department, Royal Botanic Gardens,
Kew) flying out to join Moroccan colleagues in Rabat. A tour took place along the north
coast, east to Al Hodeima before returning to Rabat. We were joined by Taleb Abdelbakir
and Yacoub from the Institut Agronomique et Veterinaire, Hassan II, Rabat. The English
trio then set out again for Taza, Oujda, the Beni Snassen Mountains and the Nador area,
but this time with Mohamed Ait Lafkih and Mouks Hassan. (Mohamed spent three
months in Banbury learning English at the ELT school in Banbury before coming to work
and train at Reading for nine further months, sponsored by the British Council. Hassan is
a graduate of Marrakech who is at present in Rabat learning English and hopes to come to
Reading to read for our MSc in plant taxonomy. Hassan became known to all as `Hassan
offendi'. Both he and Mohamed are excellent workers with a very great knowledge of
plants). Unfortunately, Dr Jury had to return to Reading for several other commitments
and flew back early from Oujda, on the Algerian border. The team continued the splendid
work and made a collection of around 500 numbers, to add to the 500 made earlier,
further west. All were collected in sets of four, (as far as possible - for Rabat, Reading,
Sevilla & Barcelona; though specimens have been divided according to which centre is
responsible for the family concerned).
Another expedition took place in June when Dr Jury joined Mohamed Ait Lafkih and
Bushra Tahiri for a week collecting around Jbel Zerhoun and Moulay-Idriss, Region 9 of
the project. An interesting collection was made around the ruins of the Roman city of
Volubilis, thanks to the co-operation of the custodian and guardians, who were,
understandably, at first a little wary of our activities! The limestone slopes and cliffs
above the cultivated areas along the Moulay-Idriss to Nzala-des-Beni-Ammar road
proved especially interesting. However, throughout the expeditions, careful attention was

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also paid to the weed and ruderal floras. This iter also yielded another 500 numbers,
again pressed into two sets of plant presses, and although there was extra work in the
field, not needing to take the specimens destined for Rabat back to Reading, the excess
baggage charges were considerably reduced.
Further expeditions are planned this autumn, continuing the successful ones last year, and
again concentrating on the abundant autumn bulbous species. It is also planned that Dr
Stephen Jury will attend the National Seminar on Biodiversity: "Resources
Phytogenetiques et Developpement Durable" to be held at the IAV, Hassan II, 24-26
October. (Professor Heywood will be speaking on "Aspects economiques de la
conservation de la biodiversite". Other delegates will be coming from Geneva, Rome,
Aleppo, Alger, Tunis, Izmir and Tambacounda).
After the March collecting expedition in northern Morocco, Mr Tim Upson remained in
Rabat, not only to ascertain that the `Restaurant Oasis' really was the best Moroccan
restaurant in the capital, but to await the end of Ramadan and the return of Dr Jury. They
were then splendidly entertained to luncheon by Mr Mohamed Ait Lafkih in his home. It
was an honour to join in the traditional feast of Eid Al-Fitir, but it was not to last as they
were joined by Bushra Tahiri and a Renault 4 belonging to the Institut Agronomique et
Veterinaire to set off on a Thymus and Lavandula hunt in the south. (Bushra Rahiri was
undertaking a PhD in Rabat on the genus Thymus; Tim's interest in Lavenders is now
It was not long before Hassan Offendi's town of Marrakech was reached and, although
we kept our eyes open for him, he had presumably just eaten so much that he was resting
indoors ... . Our usual hotel in Asni, the Grand Hotel de Toubkal, was closed for
restoration and a night was spent nearby in Ouirgane. (Probably a mistake as apparently
there is now good accommodation in Ait Lafkih's well-known village of Moulay Brahim,
near Asni. A previous expedition had visited here to shop in the souk, meet his farmer-
brother, drink tea, enjoy a tagine, as well as collect.) The following morning got off to a
good start with Lavandula maroccana observed frequently along the route, as were
several conspicuous thymes. Only brief respects were paid to the Tizi-n-Test Pass and the
Argan forests, as the night was to be spent back on the coast at Agadir. The punishing
schedule then continued to the Immouzzer Valley and Cascades (now almost completely
dried up). Again L. maroccana was frequently observed, along with many Moroccan
specialities, including the succulent Euphorbia officinarum infected by Striga
gesnerioides. Problems that arose with a puncture (and spare wheel that did not fit) were
resolved by the lighthouse keeper at Cap Rhir who happened to be a mechanic with a
well-equipped workshop, although the loss of time was regretted. The journey continued
to Tiznit and Taroudannt where good tall specimens of a Caralluma were seen. After
Taroudannt things really hotted up with Lavandulae coronopifolia, mairei, multifida and
pedunculata subsp. atlantica, all growing within a few feet of each other. (Tim used a

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complete roll of film here). A visit to Tata yielded good bushes of L. coronopifolia and L.
mairei, besides material of such desert plants as Calotropis procera, Eremophyton
chevallieri and Reboudia pinnata. The Gorges du Dades and Gorges du Todra both
provided Lavandula mairei again and Othonnopsis maroccana. For Stephen, the
highlight was in the sandy areas around Goulmima where the earlier rains meant that the
desert was in full spectacular bloom, (so very different from previous visits). The
umbellifer, Ammodaucus leucotrichus, was locally abundant with some plants almost 30
cm tall. This enigmatic genus is related to Cuminum and was studied extensively at
Reading in the 1970s (together with the rest of the tribe Caucalideae). A large collection
of desert plants was made here until it got dusk. The return journey was via Midelt and
Azrou where further collections were made along the route. Raffenaldia primuloides, an
acaulescent crucifer with spectacular yellow flowers, was seen near the Col du Zad and S.
of Azrou, near Jbel Hebri. Yet more lavenders and thymes were seen, although L.
brevidens was not seen in a recorded locality "Oued Moulouya pres de Midelt", but
further south.
The specimens were collected into pairs of presses, so the Thymes and half the other
samples could be left in Rabat. The Reading set has now been poisoned and mounted and
identification is proceeding apace, together with a gift of c.80 duplicates from John
Edmondson (LIV), also mainly from southern Morocco.
The usual three-week collecting expedition took place during the last week of April and
the first two of May, this year going to western Spain and eastern Portugal. It was jointly
organized by botanists from Salamanca (Professors Amich and Rico) and Coimbra
(Professor Jorge Paiva). Reading was represented at the `senior participant level' by Mr
Ronald Rutherford. As usual, the event was a great success, with a large number of
specimens (c.1700 gatherings) being collected from some underexplored areas. Ronnie
reports on the discovery of Drosophyllum lusitanicum in a new area. He has some
splendid slides of plants and people. The actual specimens will arrive shortly after they
have been processed in Salamanca. Several populations of European bird species were
also seen which excited the ornithologists in the party.
Next year's Iter will be in Greece and will be organized by the Flora Hellenica group.
Dr Syed Husain led a field expedition to N. Pakistan, July- August, accompanied by L.S.
Springate from the Royal Botanic Garden, Edinburgh and R.A.W. Lowe, a second year
student of our department and formerly with the NCCPG (National Council for the

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Conservation of Plants and Gardens). Dr M. Shah, a taxonomist at the Pakistan Science
Foundation also took part.
Over 700 plants were collected, including lichens, mosses and seeds from Gilgit, Yasin,
Hunza and Skardu Valleys, Khunjerab Top, and Deosai Plains.
They are grateful to the Royal Geographical Society, London for approval of the
expedition plans and for the financial help given by The Alpine Garden Society, The
Round Table Trust, The Lindeth Charitable Trust, The Percy Sladen Trust, The Oleg
Polunin Memorial Fund and the S.H. Abdeali Ziaudin Trust. They also acknowledge and
appreciate the collaboration of the Pakistan Science Foundation, Islamabad, for providing
transport and other local help.
The Association pour l'Etude Taxonomique de la Flore d'Afrique Tropicale
(A.E.T.F.A.T.) recently held a meeting in the Netherlands. Reading students and
graduates were very much in evidence at this meeting! Estrela Figueiredo, Smart Lungu,
Paul Mbugua and Francis Gachathi attended from PSL, and all presented posters.
Professor Vernon Heywood was chairman for one of the symposia.
It was reported that the hotel was in the middle of the countryside with splendid views,
but with an appalling smell of the local cows coming through the air-conditioning!
Dr Jury recently visited Trinity College, Dublin to give a well- attended seminar on his
work on the Moroccan flora. This was part of a series held weekly throughout the term.
In the evening he gave another lecture on Morocco to a joint meeting of the Irish Garden
Plant Society and the Alpine Garden Society. This concentrated on conifers and autumn-
flowering species, with special reference to conservation issues.
Stephen was also fortunate in being able to tour the National Botanic Gardens, Glasnevin
and see several splendid developments. The restored Curvilinear Glasshouse range now
towers above the surrounding builders' hoardings.. (The restoration has made use of the
ironwork of Kew's old Palm House, since this was by the same man, Richard Turner, an
Ironfounder who operated from the Hammersmith Iron Works at Ballsbridge, Dublin.
Unlike Kew, cream paint has been used (pure white being unknown in the mid-
nineteenth century) and looks aesthetically very pleasing in its garden setting. The
recreation of a piece of Burren limestone is also particularly successful. (This was
featured in one of the recent TV programmes `Over the Garden Wall' where Charles

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Nelson mentioned it was re-cycled limestone from County Laoise, but not that it came
from a cemetery!) However, Stephen found the most splendid and memorable part of his
visit to the Emerald Isle was the Old Schoolhouse at Swords!
During August and early September, James Compton visited three western States of the
USA to search for Cimicifugas, two narrowly distributed species and one of slightly
wider range. Oak Creek and a few other deep canyons in N. Arizona possess a specialised
mesophytic flora not characteristic of the nearby arid desert. Carved by streams through
hundreds of feet of red sandstone from the Coconino plateau (7000 ft), the canyon
bottoms (5000-6000 ft) are a haven for the relict species C. arizonica S. Wats. The
constantly shady, leaf-rich soil beside the Oak creek encourages such plants as Cornus
stolonifera, Rosa arizonica, Lobelia cardinalis and Polemonium albiflorum. Cimicifuga
arizonica is a perennial growing to almost 2m and producing upright virgate white-
flowered panicles covered in short glandular hairs.
Cimicifuga laciniata S. Wats. was found growing 900 miles NW of Flagstaff, Arizona, at
4000 ft, beneath Mt. Hood in Oregon State. It grows with the `Skunk Cabbage',
Lysichitum americanum, in boggy ground in deciduous streamside groves of Alnus rubra.
The plants were observed to possess short stolons, a character not known to occur in
other species within the genus. The loose cream-coloured panicles were borne 2-2.5m
above the laciniately triternate compound leaves. The third western N. American species,
Cimicifuga elata Nutt. was found growing in mixed woodland near the Columbia River
bordering Washington State; it possesses loose, small-flowered inflorescences and
biternate leaves with large palmate leaflets.
This was the usual success and again a mix of final-year students and `botanically-
minded tourists', but this year with a good contingent of enthusiastic students from the
Diploma in Scientific Horticulture course at the Royal Botanic Gardens, Kew. This happy
melange of like-minded plant scientists worked well and is something we hope we are
able to repeat next year. (The Reading party are still able to remember some of the
Crucifers!). Next year's course will take place from 23 March until 6 April (subject to
flight availability).

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Reading readers may be interested in this comprehensive and very flexible bibliographic
database system. Copies of the program are available from our Computer Services
Centre. Version 7.0.1B can be obtained for 25(pounds) and full manuals are available.
Antonio Toscano de Brito recently submitted his thesis (Systematic studies in the
subtribe Ornithocephalinae (Orchidaceae)) and has been examined by Dr Peter Gibbs; it
has been recommended to the University Senate that the degree of Doctor of Philosophy
be awarded. Antonio.
Congratulations to Dr Paul F. Cannon (BSc 1976, PhD 1981) on his promotion to
Principal Scientific Officer at the International Mycological Institute, Egham.
Readers will be delighted to learn that Dr Riyad Haddad (PhD 1987) was promoted last
January to Director of Scientific Affairs and Postgraduate Studies of the University of
Basrah. (Riyad has only Vice-Chancellor and Chancellor to go). I am envious of him
having his own private secretary and chauffeur-driven car. He is still giving lectures in
taxonomy. We look forward to the day when he can visit us again.
Khadija Biari (UNEP Course, 1993) celebrated her wedding last year and in addition to
her work at the Jardins Exotiques is engaged in teaching plant and flower culture in
Lawrence Springate (MSc, 1992) has now left Reading and taken up a job at the Royal
Botanic Garden, Edinburgh. Lawrence will be working on the Flora of Bhutan project.
Congratulations to Dr Domingo A Madulid (PhD 1980) of the Plants Unit, National
Museum, P. Burgos St., Manila, P.O. Box 2659, Philippines for securing a 3-year grant
from the John D. and Catheryn T. MacArthur Foundation based in Chicago to establish a
Biodiversity Information Centre - Plants Unit in the Philippines. The other Herbarium
News (14 (3 & 4): 12) from MO reports:
"This project is set up to serve as an information center
[sic] pertaining to plant conservation in the country. A
comprehensive database and a mini-library on plant
conservation, rare and endangered Philippine plants,
biodiversity, taxonomy, systematics and related subjects, will
be set up to serve the needs of interested scientists,
environmental planners, conservation organizations, government
agencies, students, teachers, and the public in general.
Training seminars on practical plant conservation techniques
will also be undertaken by the Unit.
The Unit will also publish pamphlets and popular articles on
rare and endangered plants, plant conservation, etc. for
dissemination to the public.

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The new unit is headed by Dr. Domingo A. Madulid with a
support staff. Any inquiries about this project can be
addressed to Dr. Domingo A. Madulid."
Congratulations to Dr Andre Mauricio de Carvalho (PhD 1989) on the receipt of The
Rupert Barneby Award. Andre, still at Centro de Pesquisas do Cacau, Cepec, Divisao de
Botanica, Caixa Postal 7, 45600 - Itabuna, Bahia, Brasil, will use his award to continue
his work on a taxonomic treatment of New World Dalbergia.
Andre shared the 1993 award with Professor Ricardo Vanni of the Instituto de Bot nica
de Nordesta, Corrientes, Argentina.
Carlos Sombrero
Carlos Sombrero (Diploma in Scientific Horticulture, Royal Botanic Gardens, Kew,
1984), BSc Botany (1987) and PhD (1992) has married Andrea Guichard in the Parish
Church of Corfe Mullen, Dorset on Saturday, 3 September. We send Dr and Mrs
Sombrero every good wish for their future together.
Dr Jim Ross (Carlos's sometime supervisor) was best man. Though the weather was not
of the best, it in no way interfered with a warm and joyous occasion. The happy couple
spent their honeymoon in the Canary Islands. The Sombreros thank the members of Plant
Science Laboratories who contributed to a cheque.
Sally has also recently submitted her PhD `Predacious and parasitoidal fungi in
association with herbivore dung in deciduous woodlands' and was examined by Dr R.C.
Cooke of Sheffield. Senate has been recommended that the degree be awarded.
We also congratulate Sally following her interview in Brussels last summer. She has been
awarded a Science and Technology Agency Fellowship Award, Programme of Japan. We
send Sally our very best wishes for her work in the Far East.
It has been recommended to Senate that Aleck be awarded the degree of Doctor of
Philosophy for his thesis on Clematis section Viorna sensu Prantl. It seems that the
External Examiner, Dr David Sutton, grows quite a number in his garden!
Congratulations Aleck.

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A card has been received from Catherine and Louis Ronse de Craene who are very happy
to announce the birth of Alexandre, in Bruges on 23 June, a brother for Camille.
Louis now has a research grant from the National Science Foundation and continues his
work on floral ontogeny. (Clearly, yet another box file for "R" reprints will be needed!)
We must also give Rita Farrell our best wishes and congratulations on her recent
Congratulations to Phil on becoming Professor of Plant Science at the Victoria University
of Wellington, PO Box 600, Wellington, New Zealand (Phone: +64 4 4721000; Fax +64
4 4715331; Internet email: Phil has now found a
house, if still haggling over price and conditions of contract. He reports:
"I have settled into the new job now, and the School of Biological Sciences has been very
welcoming. Getting on email has been an ongoing frustration for me and the network
staff, but yesterday they tracked it down to a faulty connection at the transceiver. That's
now fixed with a piece of cardboard jammed into it until a replacement part can be
obtained, and I feel like I've come alive again after being in hibernation! My research
here will be much the same, i.e. taxonomy and phylogeny of the Hebe complex, but with
an increasing amount of molecular techniques and moving also into similar work with
Ourisia. There is quite a strong group of molecular systematists here centred around
Geoff Chambers, who works mostly with birds. Currently there are student PhD
dissertations on insects, birds, reptiles, and pinnipeds, with one just starting on
Scleranthus. My teaching will be varied, but I will have quite a bit of 1st year physiology
and cell biology because those are the courses of Ken Heyes who I've replaced. I'll also
teach the 4th year course in plant systematics and maybe parts of the 3rd year course".
Best wishes from all your friends in Plant Science Laboratories, Phil. How many years
before you can come for another sabbatical here?!
The processing of the collections made by the British Bryological Society's Expedition to
Mount Mulange in 1991 continues.
This summer, we have been fortunate in having our third-year student, Victoria Bonham,
working on the family Meteoriaceae (beautiful pendant `Meteor-like' mosses, epiphytic

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on trees) for July and August. Victoria's work confirms the presence of Squamidium
brasiliense (Hornsch.) Broth., a new genus for the Malawian flora.
The Society is hoping to stage another expedition, this time to the Ruwenzori Mountains,
probably in 1996.
Dr Royce Longton has just returned from Zurich where, in his capacity as Vice-
President of the International Association of Bryologists, he officially opened the second
symposium on endangered bryophytes held 4-8 September.
Royce also gave a paper on the genetics of rare moss species in the first session on the
scientific basis of bryophyte conservation.
The above Convention was held at the University of Reading from Thursday 21st to
Sunday 25th September. The venue for lectures was the Palmer Building. On Friday
evening, around 1800 hours, the 200-odd delegates paid a visit to the School of Plant
Sciences for an informal reception in the Foyer. The place was full to capacity as
Professor Parker gave an informal welcome speech. He touched on many points of
interest to the members. He also gave his own story on how he was converted to become
an amateur in succulent plant cultivation (after a visit to Mexico where he appreciated the
cacti in their natural habitats). He outlined the wide curriculum of the School - from DNA
sequencing to landscape designs, back to the farm for basic research and over to
phytochemical issues. Several other members of PSL attended including Ronnie
Rutherford and Sue Mott (serving the drinks), Professor Caligari and Dr Terry Hedderson
(Terry even managed to find a Canadian amidst all that crowd!) Paul Mbugua had
displayed two Sansevieria posters - the genus he is researching for his PhD.
The scientific programme of lectures went well with the invited guest speakers (Charles
Glass, USA; Chuck Hanson, USA; Emst Van Jaarsveld, S. Africa; Woody Minnich,
USA; Wemer Rauh, Germany; Graham Williamson, S. Africa) each giving two lectures.
Saturday saw the arrival of the other day visitors for the shows and exhibition prepared
by Gordon Rowley and Peter Bent. One of the highlights was the auction of plants run by
Charlie Glass and Chuck Hanson. This raised over £1300 with most for the conservation
fund set up by Gordon Rowley to support projects involving the practical conservation of
the rare species. The local branch of the British Cactus and Succulent Society did an
admirable job with stewarding, airport taxi services, etc. All in all, a very popular event

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(except perhaps the queues for the food!) and a convention we hope to be repeated in
Reading in 1998!
Anybody interested in membership of the British Cactus and Succulent Society should
contact the Membership Secretary, P. Lewis, "Firgrove", 1 Springwoods, Courtmoor,
Fleet, Hampshire, GU13 9SU.
Forty members of this Society came to Reading in April for their AGM when they re-
elected Alastair Culham as editor of the Carnivorous Plant Society Journal. They
returned on Saturday, 17 September for a general meeting when five of the Society
Committee Members demonstrated propagation methods for all the main genera of
carnivorous plants. The large class all went home with plastic bags of propagation
This EU Network is co-ordinated by Professor Fred Schram, Professor of Zoology at the
University of Amsterdam. There are no less than 17 nodes in the Network in nine
countries (Denmark, France, Germany, Ireland, Italy, The Netherlands, Spain, Sweden
and the UK).
Recently Dr Jury attended a meeting in Trinity College, Dublin, hosted by Dr John
Parnell and Dr Stephen Waldren, co-ordinating this network.
Mr Tim Upson has recently attended a one-month intensive taxonomy training course in
Amsterdam, a programme organised by the Network.
The Network also has short three-month taxonomic projects in the nodes of the network -
enquiries welcome!
BAKER, W., 1994. Three men and an Orchid. Quarterly Bulletin
of the Alpine Garden Society, 62(1): 99-114; 62(2): 181-199.
COMPTON, J., 1994. 241. Salvia darcyi. The Kew Magazine,
11(2): 52-55 & pl. 241.
CULHAM, A., 1993. Sarracenia: a need for captive breeding? The
Carnivorous Plant Society Journal, 17: 5-6.

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CULHAM, A., 1993. Field notes on Borneo Nepenthes. The
Carnivorous Plant Society Journal, 17: 12-18, 20-22.
CULHAM, A., 1993. The pygmy sundew controversy: settled at last?
The carnivorous Plant Society Journal, 17: 35-38.
CULHAM, A. & GORNALL, R.J., 1994. The taxonomic significance of
napthoquinones in the Droseraceae. Biochemical Systematics
and Evolution, 22(5): 507-515.
DAVIS, A.P. & BRICKELL, C.D., 1994. Galanthus peshmenii: a new
snowdrop from the eastern Aegean. The New Plantsman, 1(1):
FIGUEIREDO, E., 1994. New records for the flora of S. Tomé and
Principe. Gascia de Orta, Sér. Bot., Lisboa, 12(1-2): 125-
FIGUEIREDO, E., 1994. Little known endemics collected by J.
Espírite Santo in S. Tomé. Garcia de Orta, Sér. Bot., Lisboa,
12(1-2): 121-124.
FIGUEIREDO, E., 1994. Contribution towards a botanical
literature for the islands of the Gulf of Guinea. Fontqueria,
34: 1-8.
Variations in the lipophilic and vacuolar flavonoids of the
genus Vellozia. Phytochemistry, 35(6): 1475-80.
HEYWOOD, V.H., 1994. Peter Hadland Davis (1918-1992). Watsonia,
20: 167-168.
JURY, S.L., 1994. Narcissus calcicola. In J. Fernández Casas
(Ed.), Asientos para una Flora Occidental, 10. Fontqueria,
39: 475.
LINNEGAR, S., 1993. The use of Iris past and present. The 1993
Iris Year Book, London: The British Iris Society.
LONGTON, R.E., 1993. British Bryological Society Expedition to
Mulanje Mountain, Malawi: 1. Background, itinerary and
procedures. Journal of Bryology, 17: 633-644.
MARHOLD, K. & RAYNER, T.G.J., 1994. Typification of the names
of two species of the Cardamine pratensis group
(Brassicaceae). Taxon, 43: 77-83.
MBUGUA, P., 1994. Three Sansevieria species of Kenya: 1.
Sansevieria robusta N.E. Br. Cactus and Succulent Journal
(US), 66: 87-88.
RAJPUT, M.T.M., TAHIR, S.S. & HUSAIN, S.Z., 1994. Taxonomic
notes on Sibbaldia micropetala (D. Don) Handel-Mazzetti.
Rosaceae. II. Candollea, 49: 129-132.

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RAJPUT, M.T.M., TAHIR, S.S. & HUSAIN, S.Z., 1994. The
distribution and synonymy of Sibbaldia procumbens L. (type
species of the genus Sibbaldia) Rosaceae. III. Candollea,
Differences in flavonoid patterns between genera within the
Velloziaceae. Phytochemistry, 36: 931-940.
YANG, Aleck Tsung-Yu, 1994. Abutilon. In Huang, Tseng-Chieng
Flora of Taiwan, second edition, 3: 739-741. Taipei, Taiwan:
National Science Council of the Republic of China.
Flora of Madeira. Edited by J.R. Press and M.J. Short. Pp.xvii + 574. The Natural History
Museum and HMSO, London, 1994. Price £35 (ISBN 0-11-310017-5).
The archipelagos comprising Macaronesia, from the Azores in the north to the Cape
Verde islands in the south, have long been of considerable interest to evolutionists and
phytogeographers. Lying along the eastern margin of the Atlantic Ocean but isolated
from the adjacent parts of southern Europe and western Africa they still retain fragments
of the Tertiary vegetation which originally flanked the Tethys Sea separating Europe
from Africa. Elements of this flora are preserved in parts of Macaronesia, notably the
Canary Islands and Madeira, their closest relatives now being found in S.E. Asia. The
subtropical rainforest (laurisilva) where these relicts now occur seems to exist because of
the mild, humid conditions in parts of these oceanic, macaronesian islands. Furthermore,
the islands' isolation has permitted evolution in plant-groups away from pressures present
in the continental areas. This had led apparently old, woody groups, such as
Argyranthemum, Echium and Sonchus, with herbaceous, often weedy representatives
elsewhere, to give rise to many endemic species in Macaronesia. This speciation within
groups which would otherwise be considered as palaeo-endemics is known as `active
epibionty' and contributes to the great scientific interest of such oceanic island-groups.
Because of their interesting and picturesque topographical diversity, their accessibility
from ± adjacent highly populated areas and their suitability for cultivation, many parts of
Macaronesia have been greatly modified by European man since their colonization in the
16th century. Nowhere has this been more profound than in the Canary Islands and
Madeira, much visited by tourists - especially from NW Europe.
The scientific, horticultural and conservation importance of such islands has been widely
recognized, as has the great value of documenting the plants which are to be found there.
This definitive Flora of Madeira fulfils the necessary criteria.
The Introduction, outlining the topography, geology, climate, vegetation and history is
accompanied by useful maps indicating features of interest. This is followed by a section

Page 17
detailing the use of the Flora and criteria followed in its preparation. A key to the
families involved then leads on to the taxonomic `meat' of the book.
The sequence of families follows that in Flora Europaea, while those of lower categories
reflect the opinions of the relevant authors. The salient features of families (except in the
pteridophytes where families are not recognized) and genera are given, whilst workable
dichotomous keys are provided for genera and the species within them. The species
descriptions are concise (again apparently owing much to Flora Europaea) but
informative, their consistency being testament to the editors' skill in dealing with
contributions from the 17 authors. In addition to the 1226 native species, of which 10%
are endemic, many naturalized taxa are described, while there are notes on species of
doubtful occurrence. The use of `like, but' or `said to occur' surely leads the way to future
theses. Ecological and geographical data are uniformly summarized, as is information on
conservation status. Original line-drawings, mostly of habit, for a significant proportion
of the endemics, together with indices to scientific and Portuguese names, complete the
This well-produced Flora is an excellent contribution to knowledge of the plants of
Macaronesia but is especially, of course, important to all drawn to the scientific and
horticultural wealth of Madeira and its adjacent islets. D.M. Moore
David Allerdice Webb
The recent tragic death of Professor David Webb, `D.A.', on the road in Oxfordshire on
September 26, cannot pass without an appreciation in Herbarium News. His numerous
visits to Reading (and he was on his way to RNG when he died) were all delightfully
memorable, if very hard work.
David was born in Dublin on August 12, 1912, educated at Charterhouse and Trinity
College, Dublin where he graduated four years later in Natural Sciences. He entered the
research world as a zoologist working on the minor elements in marine invertebrates. The
award of an 1851 "Exhibition" enabled him to transfer to Trinity College, Cambridge,
where he gained his PhD. It was during this time he had the opportunity to study in
Naples at the Marine laboratory, and gain his special affection for Italy and a love of the
Mediterranean. David returned to Dublin as an assistant to Professor H.H. Dixon, whom
he succeeded as Professor in 1950. This promotion clearly recognised his very great
abilities and his distinction in research work was further recognised by his admission to
the degree of ScD in Trinity College Dublin in 1951 and the award of an honorary
doctorate by the University of Stirling in 1979.

Page 18
David Webb retired from his post as Head of the School of Botany, Trinity College,
Dublin then as a member of their teaching staff and finally in 1984 as Curator of the TCD
Herbarium News No. 3 of 1 February 1984 reported the visit of Drs John Akeroyd and
Stephen Jury to Trinity for a symposium to mark this retirement and that an "Irish source
was quoted as saying that Professor Webb gives as many last concerts as the Rolling
Stones"! John spoke on "The future of Flora Europaea" on which he was actively
working to produce the second edition of volume one. It was this revision work which
brought David to Reading for arduous working sessions with John. David applied his
great knowledge of geography and editorial skills to carefully and accurately construct all
the geographical phrases, produce new accounts of Saxifraga and Polygonum (amongst
others), as well as place a restraining hand on John, making sure the concept of the Flora
Europaea `overview' was maintained!
These sojourns (usually over a weekend) often involved a lunch in the herbarium (Safety
Office, please proceed to the next paragraph). These `herbarium lunches', as they became
known, were no ordinary affairs, with such delights as Coquille Saint- Jacques en croute,
just for an entrée. David always arrived with a contribution to his food, sometimes a large
piece of smoked salmon, once a fresh pineapple and a pint of cream and on another
occasion, a brace of pheasants.
John always cooked an evening meal in his rooms (chez Françoise and Ian Richardson in
Whiteknights Road). John often succeeded in finding a new bird or animal previously
uneaten by DAW, the most successful being a brace of teal purchased in `Smelly Alley'.
His great generosity to enthusiastic students was boundless. Often, wined, dined and
taken on tours of Irish botanical sites, they were also treated to his wit and wisdom,
something that many Reading graduates will remember for the rest of their lives.
We are sad to report that Cheryl Langridge (BSc, Botany 1993) died recently and to
remember her University life, her family joined us to plant a tree (Betula pendula
`Tristis', a rare weeping birch) in the Harris Garden.
Oxford University Herbaria regret to announce the death of F. White, M.A., Sc.D
(Cantab.), Curator (1964-1992) and distinguished Research Curator (1992-1994).

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Terry Hedderson has been appointed an editor of Lindbergia, the journal of the Nordic
Bryological Society.
Royce Longton is now the editor of Advances in Bryology and an associate editor of the
Journal of Bryology, the journal of the British Brylogical Society. (Royce is actually
President of the British Bryological Society until December 1995).
Prof Michael W. Dick and Dr Stephen L. Jury continue to serve as Associate Editors
of The Botanical Journal of the Linnean Society, Professor Jeffrey Harborne continues
as Editor in Chief of Phytochemistry and Dr Christine Williams as an associate editor.
Dr Stephen Jury also serves on the Editorial Board of OPTIMA, responsible for
publishing the journal Flora Mediterranea. Professor Moore continues on the editorial
boards of Webbia and Parodiana.
The staff of the Department of Botany joined together for luncheon in the Orange Room
on Thursday, 12 May with Dr and Mrs E.V. Watson to celebrate Eric's 80th birthday the
following weekend. Dr Watson was presented with a copy of the recent Flora of the
Mediterranean by Christopher Grey-Wilson and was in fine form, reminiscing about the
Department in the earlier days.
Dr Renée Grayer has sadly left us for a senior post in the Jodrell Laboratories at the
Royal Botanic Gardens, Kew. She has been associated with the Department since 1976
when she came to work on her doctoral thesis on chemosystematics of Veronica L. and
related genera (partly supervised by Professor Harborne). More recently, Renée has been
working as a post-doctoral fellow on the phytoalexins of rice.
Last June we held a tea party to say goodbye, present a leaving present and wish her all
the best in her new job. We are pleased to say her new post brings her back occasionally
to Reading to develop and liaise on research projects.
We are pleased to announce that Renée's old post has been taken by Dr Vivian Dillon
who has crossed the road from the Department of Food Science to join us. Vivian
obtained her PhD in food microbiology from the University of Bath, after having already
graduated at Reading. We hope she will be very happy working with us.

Page 20
Dr Byoung-Hee Choi, a Royal Society Developing Country Fellow, has now returned
home to the Department of Biology, College of Science, Inha University, Incheon 401-
751, South Korea after spending 12 months working at the Royal Botanic Gardens, Kew
and the University of Reading.
At Reading Dr Choi learnt DNA extraction and amplification, restriction site analysis and
DNA sequencing methods. Dr Choi will be reconstructing the phylogeny of the genus on
the basis of the morphological, biochemical and molecular data using cladistic analysis.
A splendid event recently took place at the Natural History Museum: the launch of the
first published volume (six) of Flora Mesoamericana. The event started with an
introduction by The Museum's Director, Dr Neil Chalmers and an explanation of the
project by Dr Sandy Knapp. Peter Raven, who flew over especially for the event from the
Missouri Botanical Garden, then addressed the distinguished audience. It was perhaps
unfortunate that his clarion call for biodiversity was preached to the converted. This talk
was followed by wine and canapés.
The volume, written in Spanish, contains 28 families of monocotyledons (Alismataceae -
Cyperaceae), including a total of 1891 species. It can be purchased for $75 plus $3 p & p
from the Missouri Botanical Garden.
Dr Matthew Jebb of Trinity College, Dublin visited Reading to discuss research on with
Alastair Culham.
Matthew Jebb's work on Nepenthes started with his long term stay at the Christensen
Research Institute in Papua New guinea. This work resulted in publication of "An
account of Nepenthes in New Guinea" (Science in New Guinea, 17(1): 7-54 [1991]) and
will be followed by an account of Nepenthes for Flora Malesiana written jointly with
Martin Cheek at Kew.
As a result of Matthew's visit, collaborative work on Nepenthes has started.
Dr Simon Charlesworth of Downderry Nursery, Aylesford, Kent visited to consult our
Rosmarinus and Lavandula collections and to discuss their taxonomy. Tim Upson was
able to exchange several interesting taxa, which are valuable additions to our collection.

Page 21
This exchange was followed by a visit from Mrs Joan Head who is holder of one of the
NCCPG Lavandula Collections near Nottingham. Mrs Head also publishes the newsletter
The Lavendar Bag. Lavandula of all types are of especial interest to Mrs Head who was
pleased to obtain several rare Moroccan plants from us. In exchange she generously
donated many samples of wild collected seed from her own collection from the Iberian
Peninsula and the Canary Islands.
Madame Debbagh (Institut Agronomique et Vétérinaire Hassan II) visited us for two
weeks in September to study pollen in Moroccan Honey with Dr Michael Keith-Lucas.
Other visitors to the herbarium included: John Akeroyd (Kingston- upon-Thames), Yuri
Murinson (Jerusalem), Chris Chadwell (Slough), Pietro Mazzola (Palermo), Mark
Watson (Edinburgh), Peter Bailey (Adelaide), Laurie Springate (Edinburgh), Paul Wilkin
(Kew), Elizabeth Young (Milford-on-Sea), John Letts (Oxford), Dominique de Moulins
(London), Paul Hughes (Southampton), M.K. Sidhu (Punjab), Simon Charlesworth
(Aylesford), Chris Sanders (Nantwich), David Minnion (Pinner), Vahideh Nazeru
(Liverpool), Hugh McAllister (Liverpool) and Carol Palmer (Sheffield).